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The
growing trend of clonal forestry worldwide is due largely to higher
demand for quality wood fibre. Clonal eucalyptus forestry, for instance,
has already been developed in a number of countries including Brazil,
Venezuela, South Africa and Colombia.
The benefits of clonal forestry are plenty,
chief of which is improvement in wood quality particularly when
integrated with specific end uses. Given a six- to seven-year rotation,
these wood quality improvements are the main objectives of clonal
eucalyptus
The
implication is that mill and financial mangers can have the type
of wood desired for the process environment and the eventual paper
buyer. Mill and forest staff will have to work together to achieve
this.
Clonal
forestry refers to vegetatively reproduced material gown in plantations.
This material is normally obtained from rooted cuttings, but it
can also be from tissue culture, somatic embryogenesis or other
methods.
Organizations
need to look into several key issues before they take the plunge
into clonal forestry.
Cost Effectiveness
As
with any forest management technique, a number of steps must be
implemented successfully for a cost-effective clonal programme.
For a start, a wide range of species, provenance and progeny trials
are required as part of a tree improvement strategy to maintain
a wide genetic base. From this base only a small number of clones
will emerge into the operational planting programme, having been
identified in trials on multiple sites.
Besides trials, nursery technology, including
clonal garden management, is critical. Unlike seedling management,
coordination of clone demand by site and year requires careful planning
in order to maximize growth. This is due to higher initial cost
of clonal plantations compared to seedling plantations.
Changing the extensive clonal garden management
system to one of micro-cuttings is being done to reduce the cost
of each cutting. Field trials of clones on multiple sites are being
undertaken to identify disease resistance and the capacity of different
clones to respond to available resources such as water and nutrients.
And lastly, improvement in wood properties has
to be looked into. This includes selecting clones for higher wood
density as well as for better pulp and paper making ability. For
instance, the genetic modification of lignin in certain existing
clones is being evaluated. Such a modification may reduce digester
charge or time, thus lowering the cost of pulp.
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Managing Clonal Forestry
By Dr. Jeffery Wright
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Clonal
Forestry Management
Careful
management of several distinct aspects can ensure successful
clonal forestry. While each of these can be managed
separately, organisations with successful clonal strategies
must coordinate all of them including.
o Nursery management
o Silviculture
o Tree improvement
o Growth and yield
o Disease and insect monitoring and control
o Wood quality
Nursery
Management
Venezuela
is one country with a long and successful history of
forestry nursery management. Although initial efforts
were by government organisations, the private industry
is now more effective in nursery management, being supported
by research from domestic and international institutions.
Production
includes seedlings and clones of important industrial
species such as Eucalyptus urophylla, Eucalyptus urograndis,
Eucalyptus camaldulensis, Gmelina arborea and Pinus
caribaea var. hondurensis, as well as trials of other
introduced species, including Acacias. Native species
produced include mahogany, saqui saqui (Bombacopsis
quinata) apamate (Tabebuia rosea) and others.
A growing trend in Venezuela is to
develop clonal forestry for pine and eucalyptus. Given
climatic and plantation establishment constraints, however,
the clonal eucalyptus programme has to innovate in order
to be successful. Organisations cannot merely duplicate
techniques from other nurseries, as this will normally
result in failure.
Silviculture
Selection
of sites for eucalyptus clones is an ongoing task in
silviculture, which looks at the relationship of a forest
to its enviroment including the development, care and
reproduction of forest stands. There are certain soils
with characteristics that always produce high clonal
yields. On marginal soils, and these are frequent, a
number of factors are considered to determine whether
clones should be planted. Clonal genetic potential cannot
overcome poor soil or site preparation conditions.
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The
extent of site preparation depends on the soil conditions,
nutrient availability and physical structure and properties.
Typical site preparation consists of subsoiling to a
depth of 40cm, disking in two directions, followed by
bedding in sites with poor drainage. In subsequent rotations,
the only soil preparation is subsoiling to reduce any
compaction resulting from logging practices.
Control of weeds is critical in the
first 18-24 months following planting. A pre-emergent
is applied along top of the bed prior to planting. Two
or more chemical control sprayings are required in the
first year. In the second year only one chemical weeding
is required in most stands.
Fertilisation treatments have recently increased in
order to achieve higher plantation yields. Research
to date indicates that some sites are more responsive
then others and that genotypes utilise resources differently.
The research emphasis now is to identify soil parameters
associated with fertiliser response and the financial
return from fertilisation.
Tree
Improvement
A
viable and continuous investment in free improvement
must be made to ensure that clonal gains are not a one-time
occurrence. Some of the organisations that are best
known for cloning have no long-term tree improvement
programme in place.
In
Venezuela, Smurfit Carton e Venezuela (SCV) has had
a eucalyptus tree improvement programme since the mid-80s.
This programme is becoming more sophisticated as new
techniques become available.
The
SCV tree improvement programme includes the introduction
of new species and seed sources for testing Seed orchards,
control crossing, progeny testing and evaluation are
part of the improvement strategy for the seedling population.
This programme is the basis from which new clones will
e selected. All clones are tested in statistically designed
trials prior to their deployment.
Growth
and Yield
Clones
of eucalyptus can have 10-15% less taper than trees
growing n seedling stands. Thus, new volume equations
are required. In addition, selected clones have better
survival rates and less variation in height and diameter.
This makes the development of new growth and yield equations
an essential component of the clonal programme in order
to increase the accuracy of total wood projections.
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Information Resource: Asian Timber
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Disease
and Insect Moitoring and Control
The most common
problem in the case of insects is leaf-cutting ants. The ants are
controlled by chemical and mechanical methods. Other insect attacks
come from defoliators. The most effective controls are biological
especially those related to parasites or predators that reduce the
population of the target insects.
In
the early stages of new planting programmes for exotic species,
the trees are isolated from those insects and diseases that affect
them in their area of origin. Native pathogens require time to adapt
before they can infect exotic species, thereby giving a window of
opportunity to produce large quantities of wood. However, this opportunity
is short-lived and generates the need to develop innovative mechanisms
to manage insects and diseases. Diseases in the forest plantation
cannot be controlled by fungicide applications, though this is a
regular practice in the nursery.
Disease
control includes planting species or clones that are highly resistant
and maintaining a healthy tree status using appropriate silvicultural
methods.
The
constant monitoring of insects and diseases in commercial stands
and trials is very important to predict the arrival of new pests
and evaluate their impact on survival and final yield. Information
shared among numerous organisations worldwide is helping to reduce
the risk of growth loss due to disease and insects.
Wood and
Pulping Quality
In the case
of SCV, the main product of the eucalyptus clonal programme is pulp
for the manufacture of paper. Clones are tested for wood density
so that higher density clones can be selected. Individual clones
have also been tested for pulp yield, kappa number and paper strength
traits. One area of mutual interest is to modify lignin in tested
clones of eucalyptus to reduce the cost of pulping.
Outlook for
Global Clonal Forestry
Future clonal
programmes are expected to have an increasing element of biotechnology
while retaining traditional methods. Besides, marker-aided selection
will likely become more important, as will tissue culture for the
rapid bulking up of the best clones.
Movement
of eucalyptus clones worldwide will probably increase because DNA
fingerprinting and other techniques allow royalty payments to be
controlled. Improved stand management, including re-fertilisation,
will help increase yields on those sites where a response can be
predicted. In addition, mill requirements will be integrated with
clonal selection to leverage the integration the enterprise.
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